M.  Vences, Kosuch, J., Rödel, M. - O., Lötters, S., Channing, A., Glaw, F., and Böhme, W., ?Phylogeography of Ptychadena mascareniensis suggests transoceanic dispersal in a widespread Africa-Malagasy frog lineage?, Journal of Biogeography, vol. 31, pp. 593-601, 2004.