@article {406, title = {Cacosternum plimptoni: Geographic distribution (Ethiopia)}, journal = {Herpetological Review}, volume = {39}, number = {2}, year = {2008}, pages = {233}, author = {Zimkus, B.M.} } @article {409, title = {Distinguishing features of the sub-Saharan frog genera Arthroleptis and Phrynobatrachus: a short guide for field and museum-based researchers}, journal = {Breviora}, volume = {513}, year = {2008}, pages = {1-12}, author = {Zimkus, B.M. and Blackburn, D.C.} } @article {211, title = {A new species of Phrynobatrachus (Amphibia: Anura: Phrynobatrachidae) from north-western Guinea, West Africa.}, journal = {Zootaxa}, volume = {1815}, year = {2008}, pages = {43-50}, author = {Hillers, A. and Zimkus, B.M. and R{\"o}del, Mark-Oliver} } @article {377, title = {The California Academy of Sciences Gulf of Guinea expeditions (2001,2006) VI. A new species of Phrynobatrachus from the Gulf of Guinea islands and a reanalysis of Phrynobatrachus dispar and P. feae (Anura: Phrynobatrachidae)}, journal = {Proceedings of the California Academy of Sciences}, volume = {58}, number = {13-22}, year = {2007}, month = {Sep 28 2007}, pages = {367-385}, abstract = {

The oceanic islands of Sao Tome and Principe in the Gulf of Guinea of Africa harbor a surprising number of endemic amphibians. Two species of puddle frogs (Phrynobatrachus) have been described from these islands: Phryaobatrachus dispar (Peters, 1870) and P. feae (Boulenger, 1906). The validity of P. feae as a taxon distinct from P. dispar has been in doubt and in recent works the two have been considered synonymous. However, a detailed analysis has never been performed. We examined 175 specimens of Phrynobatrachus collected from the two islands during the 2001 and 2006 CAS Gulf of Guinea expeditions as well as two syntypes of P. feae. Consistent external morphological and osteological differences were found between specimens from different islands. Furthermore, maximum likelihood analysis of cytochrome b sequences revealed a high mean inter-island sequence divergence of 21\%, whereas intra-island distances were only around 1\%. This level of divergence indicates an ancient split, possibly predating the formation of Sao Tome. Mitochondrial DNA sequences from 12S rRNA, valine-tRNA, and 16S rRNA genes support this divergence and indicate that the P. dispar clade is sister to an East African clade of Phryuobatrachus, and not West African species, a recurrent theme with this insular amphibian fauna. Because the type localities of both currently available names are on Principe Island, the species endemic to Sao Tome is undescribed. We thus describe a new species of Phrynobatrachus, raising the current total of endemic amphibian species in the Gulf of Guinea Islands from six to seven.

}, url = {://PREV200700572041}, author = {Uyeda, J.C. and Drewes, R. C. and Zimkus, B.M.} } @article {410, title = {External morphology and osteology support the placement of Phrynobatrachus nlonakoensis Plath, Herrmann \& Bohme, 2006 within the genus Arthroleptis}, journal = {African Journal of Herpetology}, volume = {58}, number = {1}, year = {2009}, month = {Jun 2009}, pages = {36-38}, abstract = {

Examination of external morphology and osteology of type material of Phrynobatrachus nlonakoensis Plath, Herrmann, and Bohme, 2006 support a recent taxonomic change, transferring the species into the genus Arthroleptis, which was recently made without examination of specimens.

}, url = {://PREV200900511567}, author = {Zimkus, B.M. and Larson, J.G.} } @article {408, title = {Biogeographic analysis of puddle frogs across Cameroon and description of a new species of Phrynobatrachus (Anura: Phrynobatrachidae) endemic to Mount Oku, Cameroon}, journal = {Zoological Journal of the Linnean Society}, volume = {157}, number = {4}, year = {2009}, month = {Dec 2009}, pages = {795-813}, abstract = {

Patterns of species divergence amongst 13 species of Phrynobatrachus from Cameroon were analysed using phylogenetic analyses of mitochondrial DNA sequence data from 12S rRNA, valine-tRNA, and 16S rRNA. A clade endemic to the mountains of western Cameroon and eastern Nigeria, part of the geologic rift zone termed the Cameroon Volcanic Line, supports the view that montane regions have acted as centres of speciation. Bioko Island (Equatorial Guinea), located 32 km off the coast of Cameroon, is populated by three different lowland Phrynobatrachus species from the mainland. Two of these species, Phrynobatrachus auritus and Phrynobatrachus cornutus reach submontane elevations on Bioko Island (up to 1200-1300 m) but are present only at lowland elevations on mainland Cameroon. A new miniature species of puddle frog of the genus Phrynobatrachus restricted to Mt Oku in the Republic of Cameroon is diagnosed on the basis of morphological characters and molecular data. Unlike all other miniature Phrynobatrachus (snout-vent length \< 20 mm), males of this species have unique ventral coloration characterized by a black throat, dark brown to black pectoral and abdominal regions, and a light or white area on the proximal hind limbs. Minute spines cover the venter in males, often appearing as white asperities. The description of this new species supports the significance of this montane ecoregion as a biodiversity hotspot for anurans. (C) 2009 The Linnean Society of London, Zoological Journal of the Linnean Society, 2009, 157, 795-813.

}, url = {://PREV201000046439}, author = {Zimkus, B.M.} } @article {2307, title = {Examination of the molecular relationships of sand frogs (Anura: Pyxicephalidae: Tomopterna) and resurrection of two species from the Horn of Africa}, journal = {Zootaxa}, volume = {2933}, year = {2011}, pages = {27{\textendash}45}, author = {Zimkus, B.M. and Larson, J.G.} } @article {411, title = {Complex patterns of speciation and diversity among African frogs (genus Phrynobatrachus)}, journal = {Molecular Phylogenetics \& Evolution}, volume = {55}, year = {2010}, pages = {883-900}, author = {Zimkus, B.M. and R{\"o}del, Mark-Oliver and Hillers, A.} } @article {412, title = {Light at the end of the tunnel: insights into the molecular systematics of East African puddle frogs (Anura: Phrynobatrachidae)}, journal = {Systematics and Biodiversity}, volume = {8}, year = {2010}, pages = {39-47}, author = {Zimkus, B.M. and Schick, S.} } @article {358, title = {Systematics of {\textquoteright}Little Brown Frogs{\textquoteright} from East Africa: Recognition of Phrynobatrachus scheffleri and description of a new species from the Kakamega Forest, Kenya (Anura: Phrynobatrachidae)}, journal = {Salamandra}, volume = {46}, year = {2010}, pages = {24-36}, author = {Schick, S. and Zimkus, B.M. and Channing, A. and Koehler, J. and L{\"o}tters, S.} }